He would eventually enact a plot to transform the human race into powerful superhumans like himself, and be a god to rule over them in a new age. Before being killed by Wesker, Oswell Spencer reveals that Wesker was the survivor of a Progenitor virus variant administration experiment-the 'Wesker Project.' (It is stated in the RE5 downloadable content 'Lost in Nightmares' that another Wesker, Alex, survived as well). Wesker returns in Resident Evil 5 conspiring with the pharmaceutical division of the TRICELL conglomerate as a means to create the 'Uroboros virus', an enhanced virus derived from the Progenitor virus, with which he ultimately plans to release into the Earth's atmosphere. In the expanded release, Code: Veronica X, he briefly confronts Chris's younger sister Claire and nearly kills her in order to torment Chris, but is called away by his associates and thus chooses to spare her life. He confronts both Chris and Alexia and he took the corpse of Steve Burnside in order to extract the T-Veronica virus. When Chris Redfield knocks his sunglasses off during a brief struggle, Wesker's orange, almost reptilian eyes are seen for the first time. He orchestrates a raid on Rockfort Island in order to capture Alexia Ashford, creator of the T-Veronica virus who now holds the only remaining sample of the virus within her body. Wesker returned in Resident Evil – Code: Veronica, augmented with superhuman powers and working as an agent for 'H.C.F.' (Hive/Host Capture Force), a Special Forces unit of the rival company opposed to Umbrella. However, his plan was ruined after the Tyrant was destroyed by his former subordinates, and he survived his apparent death in the first game by injecting himself beforehand with an experimental variant of the T-virus, provided by his former Umbrella colleague William Birkin, that revived and augmented him with superhuman strength, speed and regeneration, but at the expense of his humanity. field as a means to buy his way into a high position, and he deployed the elite STARS against it in order to obtain combat data to prove the Tyrant's effectiveness to the company. In the special giveaway fictional documentary titled Wesker's Report (rewritten in 2003 to include details from Resident Evil 0 and the Resident Evil remake), Wesker reveals that he planned to sell the Tyrant to Umbrella's rival company in the B.O.W. However, this is part of an even bigger plan. The Sega Saturn port of Resident Evil features a Battle Mode minigame where the player can fight a zombified version of Wesker.

Chris Redfield, Jill Valentine and the others come to learn of Wesker's true motive, but after releasing the Tyrant monster, Wesker is seemingly killed.

Wesker is eventually revealed to be a double agent working for the Umbrella Corporation ordered by his superiors to lure the STARS into the Spencer mansion to be used as test subjects against the mutated creatures to gather battle data, although he later implies that this was itself a cover for his own agenda regarding stealing Umbrella's research.

In that game, he is the commanding officer of Alpha team of the Special Tactics And Rescue Service (STARS), where he initially helps the player by leaving supplies and useful information.

Through the course of much betrayal of his allies to further his own plans, Wesker fakes his death, gains superhuman abilities from an experimental variant of the Progenitor virus, and works with both Umbrella's mysterious rival company and their successors in the field of biological weapons development, TRICELL, until his ultimate defeat by Chris Redfield in Resident Evil 5.Wesker made his first appearance as a non-playable character in the original Resident Evil game. To this end, he is affiliated with the Umbrella Corporation as one of its most promising researchers, and at the same time participates in illicit activities by going undercover as a S.T.A.R.S captain in Raccoon City. Wesker seeks to replace humanity through mass extinction and forced evolution, believing it to be an evolutionary dead end. Introduced in the first game, Wesker has been one of the main antagonists throughout the series, where he usually manipulates story events behind-the-scenes.

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